Vegetable-fern, Paco fern

Where does Vegetable-fern grow?

Found in: Africa, Asia, Bangladesh, Bhutan, Bougainville, Cambodia, China, Fiji, Hawaii, Himalayas, India, Indochina, Indonesia, Japan, Laos, Malaysia, Nepal, Nigeria, Northeastern India, NW India, Oceania, Pacific, Papua New Guinea, PNG, Philippines, SE Asia, Sikkim, Singapore, Solomon Islands, Sri Lanka, Taiwan, Thailand, Tibet, United States, Vanuatu, Vietnam, West Africa, West Papua

Notes: There are about 400 Diplazium species. They have also been put in the families Aspleniaceae and Woodsiaceae.

Status: It is a commercially cultivated vegetable. One of the commonly used and popular ferns of the Asian tropical lowland areas. Young freshly uncurled fronds are common in markets.

Growing Vegetable-fern, Paco fern

Cultivation: Plants can be grown from spores. They need to be in a well drained potting medium and kept with a high humidity. The spores need a temperature of 21°C and sown as soon as ripe. Plants should be transplanted into a moist well drained soil with partial shade. Plants can also be grown by separating out the underground runners.

Edible Uses: The fronds are cooked and eaten as a vegetable. They are also used in stews. They can be fried. They can be steamed in earth ovens. They are cooked with daal. Very young leaves are also eaten raw in salads.

Production: The fronds will only store for a very sort time so need to be used quickly.

Nutrition Info

Vegetable-fern, Paco fern Photos

References

Alegado, A. M. & De Guzman, R. B., 2014, Indigenous food crops of the Aetas tribe in the Philippines and their traditional methods of food preparation. in Promotion of Underutilized Indigenous Food Resources for Food Security and Nutrition in Asia and Pacific. FAO. Bangkok p 160 (As Athyrium esculentum)

Ambasta S.P. (Ed.), 2000, The Useful Plants of India. CSIR India. p 60 (As Athyrium esculentum)

Amoroso, V., et al, 2017, Indigenous edible ferns in southern Philippines: Utilization and conservation. China_Botanical Congress. p 3

Amoroso, V. B., et al., 2017, Bringing back the Lost Value of Philippine EdibleFerns: Their Antioxidant, Proteins and Utilization. International Journal of Advanced research. 5(4), 757-770

Anderson, E. F., 1993, Plants and people of the Golden Triangle. Dioscorides Press. p 209

Azam, F. M. S., et al, 2014, Are Famine Food Plants Also Ethnomedicinal Plants? An Ethnomedicinal Appraisal of Famine Food Plants of Two Districts of Bangladesh. Evidence-Based Complementary and Alternative Medicine Volume 2014, Article ID 741712, 28 pages

Bandyopadhyay, S. et al, 2009, Wild edible plants of Koch Bihar district, West Bengal. Natural Products Radiance 8(1) 64-72

Bandyopadhyay, S., et al, 2012, A Census of Wild Edible Plants from Howrah District, West Bengal, India. Proceedings of UGC sponsored National Seminar 2012

BARC, 2016, State of Biodiversity for Food and Agriculture in Bangladesh. Bangladesh Agricultural Research Council.

Bhattarai, S. & Rajbhandary, S., 2017, Pteridophyte Flora of Manaslu Conservation Area, Central Nepal. American Journal of Plant Sciences, 2017, 8, 680-687

Brickell, C. (Ed.), 1999, The Royal Horticultural Society A-Z Encyclopedia of Garden Plants. Convent Garden Books. p 371

Brown, W.H., 1920, Wild Food Plants of the Philippines. Bureau of Forestry Bulletin No. 21 Manila. p 19

Burkill, I.H., 1966, A Dictionary of the Economic Products of the Malay Peninsula. Ministry of Agriculture and Cooperatives, Kuala Lumpur, Malaysia. Vol 1 (A-H) p 848

Call, C. A., et al, 2004, Participatory Rural Appraisal in the Upland Ecosystem of Mt Malindang, Misamis Occidental, Philippines. Biodiversity Research Programme for Development in Mindanao.

Cao, Y., et al, 2020, Ethnobotanical study on wild edible plants used by three trans-boundary ethnic groups in Jiangcheng County, Pu’er, Southwest China. Journal of Ethnobiology and Ethnomedicine (2020) 16:66

Chai, P. P. K. (Ed), et al, 2000, A checklist of Flora, Fauna, Food and Medicinal Plants. Lanjak Entimau Wildlife Sanctuary, Sarawak. Forestry Malaysia & ITTO. p 159

Chai, P.P.K., 2000, Development of the Lanjak Entimau Wild life sanctuary. Final report. ITTO and Sarawak Forestry. p 19

Chai, T., et al, 2015, Evaluation of Glucosidase Inhibitory and Cytotoxic Potential of Five Selected Edible and Medicinal Ferns. Tropical Journal of Pharmaceutical Research March 2015; 14 (3): 449-454.

Chao, Pi-Yu, et al, 2014, Antioxidant Activity in Extracts of 27 Indigenous Taiwanese Vegetables. Nutrients 2014, 6, 2115-2130 (As Anisogonium esculentum)

Chettri, S., et al, 2018, Nutrient and Elemental Composition of Wild Edible Ferns of the Himalaya. American Fern Journal 108(3):95–106

Chowdery, T., et al, 2014, Wild edible plants of Uttar Dinajpur District, West Bengal. Life Science Leaflets. 47:pp 20-36 http://lifesciencesleaflets.ning.com

Chowdhury, A. & Das, A. P., 2014, Conservation through sustainable utilization of wetland leafy vegetables of Terai and Duars, West Bengal, India. International Journal of Advanced Life Sciences (IJALS), 7(4) p 654

Chowdhury, M. & Mukherjee, R., 2012, Wild Edible Plants Consumed by Local Communities of Maldah of West Bengal, India. Indian J.Sci.Res.3(2) : 163-170

Dangol, D. R. et al, 2017, Wild Edible Plants in Nepal. Proceedings of 2nd National Workshop on CUAOGR, 2017.

Deb, D., et al, 2013, Wild Edible Plants and Their Utilization in Traditional Recipes of Tripura, Northeast India. Advances in Biological Research 7(5):203-211

Delos Angeles, M. D., & Buot Jr., I. E., 2012, Orders and Families of Philippine Pteridophytes. Journal of Nature Studies 11 (1 & 2): 19-33

Dobriyal, M. J. R. & Dobriyal, R., 2014, Non Wood Forest Produce an Option for Ethnic Food and Nutritional Security in India. Int. J. of Usuf. Mngt. 15(1):17-37

Facciola, S., 1998, Cornucopia 2: a Source Book of Edible Plants. Kampong Publications, p 45

Flora of Solomon Islands

Food Composition Tables for use in East Asia FAO http://www.fao.org/infoods/directory No. 502

French, B.R., 1986, Food Plants of Papua New Guinea, A Compendium. Asia Pacific Science Foundation p 84

French, B.R., 2010, Food Plants of Solomon Islands. A Compendium. Food Plants International Inc. p 106

Furusawa, T., et al, 2014, Interaction between forest biodiversity and people's used of forest resources in Roviana, Solomon Islands: implications for biocultural conservation under socioeconomic changes. Journal of Ethnobiology and Ethnomedicine, 10:10

Gangwar, A. K. & Ramakrishnan, P. S., 1990, Ethnobotanical Notes on Some Tribes of Arunachal Pradesh, Northeastern India. Economic Botany, Vol. 44, No. 1 pp. 94-105

Garcia, G. S. C., 2006, The mother-child nexus. Knowledge and valuation of wild food plants in Wayanad, Western Ghats, India. Journal of Ethnobiology and Ethnomedicine, 2:39

Ghorbani, A., et al, 2012, A comparison of the wild food plant use knowledge of ethnic minorities in Naban River Watershed Nature Reserve, Yunnan, SW China. Journal of Ethnobiology and Ethnomedicine; 8:17

Ghosh, C. & Das A. P., 2011, Some useful and poisonous tea garden weeds from the Darjiling District of West Bengal, India. Pleione 5(1): 91 - 114

Goudey, C.J., 1988, A Handbook of Ferns for Australia and New Zealand. Lothian. p 97

Hariyadi, B., 2008, The Entwined Tree: Traditional Natural Resource Management of Serampas, Jambi, Indonesia. Ph. D thesis. Univ. or Hawaii. p 444 (As Athyrium esculentum)

Hedrick, U.P., 1919, (Ed.), Sturtevant's edible plants of the world. p 279

Henderson, C.P. and I.R.Hancock, 1988, A Guide to the Useful Plants of the Solomon Islands. Res. Dept. Min of Ag. & Lands. Honiara, Solomon Islands. p 84

Hoe, V. B. & Siong, K. H., 1999, The nutritional value of indigenous fruit and vegetables in Sarawak. Asia Pacific J. Clin. Nutr. 8(1):24-31 (As Diplazium esculentum)

Hossain, U. & Rahman, A., 2018, Study and quantitative analysis of wild vegetable floral diversity available in Barisal district, Bangladesh. Asian J. Med. Biol. Res. 2018, 4 (4), 362-371

Hu, Shiu-ying, 2005, Food Plants of China. The Chinese University Press. p 273 (As Anisogonium)

Irawan, D., et al, 2006, Ethnobotanical study and nutrient potency of local traditional vegetables in Central Kalimantan. Tropics Vol. 15(4)

Islam, M., 1983, Utilisation of Certain Ferns and Fern Allies in the North-Eastern Region, India. J.Econ. Tax. Bot. Vol.4.No.3 pp 861- (As Athyrium esculentum)

IUCN (2013). Ecological Survey of the Mekong River between Louangphabang and Vientiane Cities, Lao PDR, 2011-2012. Vientiane, Lao PDR: IUCN. 241 pp.

Jacquat, C., 1990, Plants from the Markets of Thailand. D.K. Book House p 33

Japanese International Research Centre for Agricultural Sciencewww.jircas.affrc.go.jp/project/value_addition/Vegetables

J. Bot. (Schrader) 1801(1,2):312. 1803-1804

Johnson, N., 2002, Environmental Change in northern Thailand: Impact on Wild Edible Plant Availability. Ecology of Food and Nutrition, 41: 5, 373-399

Joshi, K. and Joshi, A. R., 2008, Ethnobotanical Studies on Some Lower Plants of the Central Development Region, Nepal. Ethnobotanical Leaflets 12:832-40

Joshi, N. & Siwakoti, M., 2012, Wild Vegetables Used by Local Community of Makawanpur District and Their Contribution to Food Security and Income Generation. Nepal Journal of Science and Technology Vol. 13, No. 1 (2012) 59-66

Kar, A., 2004, Common wild vegetables of Aka tribe of Arunachal Pradesh. Indian Journal of Traditional Knowledge 3(3) pp 305-313

Kar, A., et al, 2013, Wild Edible Plant Resources used by the Mizos of Mizoram, India. Kathmandu University Journal of Science, Engineering and Technology. Vol. 9, No. 1, July, 2013, 106-126

Kays, S. J., and Dias, J. C. S., 1995, Common Names of Commercially Cultivated Vegetables of the World in 15 languages. Economic Botany, Vol. 49, No. 2, pp. 115-152

Konsam, S., et al, 2016, Assessment of wild leafy vegetables traditionally consumed by the ethnic communities of Manipur, northeast India. Journal of Ethnobiology and Ethnomedicine, 12:9

Kiple, K.F. & Ornelas, K.C., (eds), 2000, The Cambridge World History of Food. CUP p 1875

Kumar, A., et al, 2012, Ethnobotanical Edible Plant Biodiversity of Lepcha Tribes. Indian Forester, 138 (9):798-803

Kumar, S. A., Manus, D. & Mallika, M., 2018, Impact of non-timber forest products on Forest and in Livelihood Economy of the People of Adjoining Areas of Jalpaiguri Forest Division, West Bengal, India. Int. J. of Life Sciences, 2018; 6 (2):365-385

Kuo, W. H. J., (Ed.) Taiwan's Ethnobotanical Database (1900-2000), http://tk.agron.ntu.edu.tw/ethnobot/DB1.htm

Lacuna-Richman, C., 2006, The use of non-wood forest products by migrants in a new settlement: experiences of a Visayan community in Palawan, Philippines. Journal of Ethnobiology and Ethnomedicine. 2:36 (As Athyrium)

Leach, G. J., 1988, Bush Food Plants of the Blackwater and Karawari Rivers Area, East Sepik Province, Papua New Guinea. Science in New Guinea 14(2). p 98

Li, S., et al, 2020, Monpa, memory, and change: an ethnobotanical study of plant use in Mêdog County, South-east Tibet, China. Journal of Ethnobiology and Ethnomedicine. (2020) 16:5 p 16

Liu, Y., et al, 2012, Food uses of ferns in China: a review. Acta Societatis Botanicorum Poloniae 84(4): 263-270 (As Callipteris esculenta) (Also as Callipteris esculenta var. pubescens)

Luczaj, L., et al, 2021, Wild food plants and fungi sold in the markets of Luang Prabang, Lao PDR. Journal of Ethnobiology and Ethnomedicine (2021) 17:6

Lugod, G.C. and de Padua L.S., 1979, Wild Food Plants in the Philippines. Vol. 1. Univ. of Philippines Los Banos. p 66

Luo, B., et al, 2019, Wild edible plants collected by Hani from terraced rice paddy agroecosystem in Honghe Prefecture, Yunnan, China. Journal of Ethnobiology and Ethnomedicine 15:56 (As Callipteris esculenta)

Macmillan, H.F. (Revised Barlow, H.S., et al), 1991, Tropical Planting and Gardening. Sixth edition. Malayan Nature Society. Kuala Lumpur. p 356 (As Asplenium esculentum)

Maghirang, R. G., et al, 2018. Ethnobotanical Studies of Some Plants Commonly Used as Vegetables in Selected Provinces of the Philippines. Journal of Nature Studies. 17(2), 30-43.

Manandhar, N.P., 2002, Plants and People of Nepal. Timber Press. Portland, Oregon. p 203

Marandi, R. R. & Britto, S. J., 2015, Medicinal Properties of Edible Weeds of Crop Fields and Wild plants Eaten by Oraon Tribals of Latehar District, Jharkhand. International Journal of Life Science and Pharma Research. Vo. 5. (2) April 2015

Maroyi, A., 2014, Not just minor wild edible forest products: consumption of pteridophytes in sub-Saharan Africa. Journal of Ethnobiology and Ethnomedicine. 10:78

Medhi, P. & Borthakur, S. K., 2012, Phytoresources from North Cachur Hills of Assam -3: Edible plants sold at Hflong market. Indian Journal or Natural Products and Resources. 3(1) pp 84-109

Medhi, P. & Borthakur, S. K., 2013, Wild edible plants sold by the Zeme Nagas at the makeshift market of Mahur, Dima Hasao district of Assam. Pleione 7(1): 84 - 93. 2013

Medhi, P., Sarma, A and Borthakur, S. K., 2014, Wild edible plants from the Dima Hasao district of Assam, India. Pleione 8(1): 133-148

Monsalud, M.R., Tongacan, A.L., Lopez, F.R., & Lagrimas, M.Q., 1966, Edible Wild Plants in Philippine Forests. Philippine Journal of Science. p 445 (As Athyrium esculentum)

Mot So Rau Dai an Duoc O Vietnam. Wild edible Vegetables. Ha Noi 1994, p 166

Narayanan Ratheesh, M. K. et al, 2011, Wild edible plants used by the Kattunaikka, Paniya and Kuruma tribes of Wayanad District, Kerala, India. Journal of Medicinal Plants Research Vol. 5(15), pp. 3520-3529

Nombo, P. & Leach, J. 2010, Reite Plants: An Ethnobotanical Study in Tok Pisin and English. ANU press. p 111 (As Athyrium)

Ochse, J. J. et al, 1931, Vegetables of the Dutch East Indies. Asher reprint. p 600

Ogle, B. M., et al, 2003, Food, Feed or Medicine: The Multiple Functions of Edible Wild Plants in Vietnam. Economic Botany 57(1): 103-117

Ong, H., et al, 2012, Traditional knowledge and usage of edible plants among the Semai community of Kampung Batu 16, Tapah, Perak, Malaysia. Scientific Research and Essays Vol. 7(4), pp. 441-445, 30 January, 2012

Oomen, H.A.P.C., & Grubben, G.J.H., 1978, Tropical Leaf Vegetables in Human Nutrition, Communication 69, Department of Agricultural research, RTI Amsterdam, p 36

Ojha, R. & Devkota, H. P., 2021, Edible and Medicinal Pteridophytes of Nepal: A Review. Ethnobotany Research & Applications 22:16

Pagag, K. & Borthakur, S.K., 2012, Wild edible wetland plants from Lakhimpur district of Assam, India. Pleione 6(2): 322 - 327

Partha, P., 2014, Ethnobotany of the Laleng (Patra) Community in Bangladesh. Journal of Pharmacognosy and Phytochemistry. 2(6):173-184

Patiri, B. & Borah, A., 2007, Wild Edible Plants of Assam. Geethaki Publishers. p 162

Pawera, L., et al, 2020, Wild Food Plants and Trends in Their Use: From Knowledge and Perceptions to Drivers of Change in West Sumatra, Indonesia, Foods. 2020, 9, 1240

Peekel, P.G., 1984, (Translation E.E.Henty), Flora of the Bismarck Archipelago for Naturalists, Division of Botany, Lae, PNG. p 13, 15.

Pegu, R., et al, 2013, Ethnobotanical study of Wild Edible Plants in Poba Reserved Forest, Assam, India. Research Journal of Agriculture and Forestry Sciences 1(3):1-10

Pfoze, N. L., et al, 2012, Assessment of Local Dependency on Selected Wild Edible Plants and fruits from Senapati district, Manipur, Northeast India. Ethnobotany Research & Applications 10:357-367

Pfoze, N. L., et al, 2012, Survey and assessment of floral diversity on wild edible plants from Senapati district of Manipur, Northeast India. Journal or Biodiversity and Environmental Sciences. 1(6):50-52

Pham-Hoang Ho, 1999, An Illustrated Flora of Vietnam. Nha Xuat Ban Tre. p 164

Phawa, G. M., Dkhar, E. K. & Marbaniang, D., 2019, Indigenous Wild Edible Plants of Bataw Village, East Jaintia Hills District, Meghalaya. International Journal of Arts, Science and Humanities. 7(2)

Piggott, A. G. 1988, Ferns of Malaysia in Colour. Tropical press, Malaysia. p 309

Priyadi, H., et al, 2010, Five hundred plant species in Gunung Halimun Salak National Park West Java. A checklist including Sundanese names, distribution and use. CIFOR, FFPRI, SLU p 125

Ramachandran, V. S., 2007, Wild edible plants of the Anamalais, Coimbatore district, western Ghats, Tamil Nadu. Indian Journal or Traditional Knowledge. 6(1) pp 173-176

Ramachandran,V. S., & Udhayavani, C., 2013, Knowledge and uses of wild edible plants by Paniyas and Kurumbas of Western Nilgiris, Tamil Nadu. Indian Journal of Natural Products and Resources. 4(4) December 2013, pp 412-418

Rao, M. L. S., et al, 2014, Indigenous Plant Foods which are commonly consumed by the tribal communities in Dumbriguda Area of Visakhapatnam District, Andhra Pradesh, India. Biolife. Vol 2, Issue 3

Rashid, A., Anand, V.K. & Serwar, J., 2008, Less Known Wild Plants Used by the Gujjar Tribe of District Rajouri, Jammu and Kashmir State. International Journal of Botany 4(2):219-244

Saikia, M., 2015, Wild edible vegetables consumed by Assamese people of Dhemaji District of Assam, NE India and their medicinal values. Archives of Applied Science Research, 2015, 7 (5):102-109

Sakar, A. & Das, A. P., 2018, The traditional knowledge on edible wild leafy vegetables of Rabha Tribe in Duars of North Bengal: a potential reinforcement to food security. Pleione 12(2): 275 - 281. 2018.

Sakunpak, A. & Panichayupakaranant, P., 2012, Antibacterial activity of Thai edible plants against gastrointestinal pathogenic bacteria and isolation of a new broad spectrum antibacterial polyisoprenylated benzophenone, chamuangone. Food Chemistry 130 (2012) 826–831

Sang, D. T., & Mizoue, K. O. N., 2012, Use of Edible Forest Plants among Indigenous Ethnic Minorities in Cat Tien Biosphere Reserve, Vietnam. Asian Journal of Biodiversity Vol. 3 (1), p 23-49

Sarker, S. K., & Enayet Hossain, A. B. M., 2009, Pteridophytes of Greater Mymensingh District of Bangladesh used as Vegetables and Medicines. Bangladesh J. Plant Taxon. 16(1) 47-56. p 52

Seal, T., 2012, Evaluation of Nutritional Potential of Wild Edible Plants, Traditionally Used by the Tribal People of Meghalaya. American Journal of Plant Nutrition and FertilizationTechnology 2(1): 19-26

Singh, K.K., & Maheshwari, J.K., 1984, Studies of the Flora of Dudwa National Park. Kheri District, U.P. (Part 1) J. Econ. Tax. Bot. Vol 5 No. 2 pp 379

Solomon, C., 2001, Encyclopedia of Asian Food. New Holland. p 144 (As Athyrium esculentum)

Srichaiwong, P., et al, 2014, A Study of the Biodiversity of Natural Food Production to Support Community Upstream of Chi Basin, Thailand. Asian Social Science 10 (2):

Srivastava, R. C., 2010, Traditional knowledge of Nyishi (Daffla) tribe of Arunachal Pradesh. Indian Journal of Traditional Knowledge. 9(1):26-37

Staples, G.W. and Herbst, D.R., 2005, A tropical Garden Flora. Bishop Museum Press, Honolulu, Hawaii. p 29

Sujarwo, W., et al, 2014, Ethnobotanical Study of Edible Ferns Used in Bali Indonesia. Asia Pacific Journal of Sustainable Agriculture Food and Energy. 2(2): 1-4.

Sukarya, D. G., (Ed.) 2013, 3,500 Plant Species of the Botanic Gardens of Indonesia. LIPI p 865

Sukenti, K., et al, 2016, Ethnobotanical study on local cuisine of the Sasak tribe in Lombok Island, Indonesia. Journal of Ethnic Foods. 3 (2016) 189-200 p 198

Sundriyal, M., et al, 1998, Wild edibles and other useful plants from the Sikkim Himalaya, India. Oecologia Montana 7:43-54

Sundriyal, Manju and R. C. Sundriyal, Wild Edible Plants of the Sikkim Himalaya: Marketing, Value Addition and Implications for Management. Economic Botany: Vol. 58, No. 2, pp. 300–315.

Sundriyal, M., et al, 2004, Dietary Use of Wild Plant Resources in the Sikkim Himalaya, India. Economic Botany 58(4) pp 626-638

Tanaka, Y. & Van Ke, N., 2007, Edible Wild Plants of Vietnam: The bountiful garden. Orchid books. p 36

Taram, M., et al, 2018, Wild Food Plant Resources of Komkar Adi Tribe of Upper Siang District in Arunachal Pradesh, India. Bulletin of Arunachal Forest Research, Vol. 33(2), 27-35

Teron, R. & Borthakur, S. K., 2016, Edible Medicines: An Exploration of Medicinal Plants in Dietary Practices of Karbi Tribal Population of Assam, Northeast India. In Mondal, N. & Sen, J.(Ed.) Nutrition and Health among tribal populations of India. p 149

Thapa, L. B., et al, 2014, Wild Edible Plants used by endangered and Indigenous Raji Tribe in Western Nepal. International Journal of Applied Sciences and Biotechnology. Vol 2(3):243-252

Thoa, P. T. K., et al, 2013, Biodiversity indices and utilization of edible wild plants a case study of the Cham Island in Quang Nam Province, Vietnam. Journal of research in Environmental Science and Toxicology Vol. 2(9): 167-174

Tsering, J., et al, 2017, Ethnobotanical appraisal on wild edible plants used by the Monpa community of Arunchal Pradesh. Indian Journal of Traditional Knowledge. Vol 16(4), October 2017, pp 626-637

Tsherig, K., 2012, Edible Wild Plants of Bhutan and their contribution to Food and Nutrition Security. Ministry of Ag. and Forests, Bhutan. www.fao.org

Turner, N. J. et al, 2011, Edible and Tended Wild Plants, Traditional Ecological Knowledge and Agroecology. Critical Reviews in Plant Sciences, 30:198-225

Uprety, Y., et al, 2012, Diversity of use and local knowledge of wild edible plant resources in Nepal. Journal of Ethnobotany and Ethnomedicine 8:16

USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network - (GRIN). [Online Database] National Germplasm Resources Laboratory, Beltsville, Maryland. Available: www.ars-grin.gov/cgi-bin/npgs/html/econ.pl (10 April 2000)

Van Sam, H. et al, 2008, Uses and Conservation of Plant Species in a National Park. A case study of Ben En, Vietnam. Economic Botany 62:574-593

Wahyudi, 2017, Non-timber Forest Product (NTFP) Commodities Harvested and Marketed by Local People at the Local Markets in Manokwari - West Papua. Indonesian Journal of Forestry Research Vol. 4, No. 1, 27-35

Wild edible plants of Himachal Pradesh (As Athyrium esculentum)

Wiriadinata, H., 1995, Ethnobotany of Economic Plants in the Baliem Valley, Jayawijaya, Irian Jaya, Indonesian Institute of Science, Bogor, Indonesia

World Checklist of Useful Plant Species 2020. Royal Botanic Gardens, Kew

Xu, You-Kai, et al, 2004, Wild Vegetable Resources and Market Survey in Xishuangbanna, Southwest China. Economic Botany. 58(4): 647-667. (As Callipteris esculenta) (Also as Callipteris esculenta var. pubescens)